Introduction to Cervids


Chapter 1
Introduction to Cervids


Sam Ecroyd, Peter Green, Kit Heawood, Mitchell Palmer and Jim Walsh


Introduction


The interactions between veterinarians and deer can be many and varied, from sporadic encounters with wild deer involved in road traffic collisions to regular herd health planning visits on deer farms. From a veterinary perspective, deer are fascinating creatures. They exploit and thrive in many different ecological niches and can be found across the world in many forms. They are remarkably hardy creatures, often dealing with injuries and diseases in ways that would be surprising in domestic species. They have the ability to grow and repair bone at a rate that is unimaginable in other animals. Yet with such an abundance of deer worldwide, they are species that are often sidelined or neglected within veterinary schools. Many veterinarians know little about them.


History


Humans and deer have interacted for millennia; early man hunted deer for food and skins and collected antlers to make tools. Reindeer (Rangifer tarandus) were domesticated as beasts of burden and as providers of food more than 5000 years ago. They have become integral to the ways of life of various human cultures. Historically, deer were widely hunted for sport by the nobility, and entire swathes of countryside were dedicated to their keeping and breeding. In more modern times, attempts have been made to domesticate other deer species, leading to the first new large animal livestock species in several thousand years. Deer have permeated our folklore, be they the ‘fairy cattle’ of Irish myth or the noble white stag predicting the reigns of Kings and Queens. In the United Kingdom, deer have given their names to towns, streets and pubs such as the White Hart and the Roebuck. Deer frequently appear in European heraldry and insignia.


Deer Species


The taxonomic family ‘Cervidae’ contains a vast range of species, from the diminutive pudu to the mighty moose. While there is more genetic variation between a roe deer and a fallow deer than between a cow and a sheep, these widely differing animals are frequently grouped together under the catch-all term ‘deer’. This can pose somewhat of a challenge to the veterinarian in general practice when faced with a ‘deer’, because different species can vary substantially in their handling and treatment. This chapter provides a brief overview of some of the species most commonly seen in farms, parks and the wild (primarily in the United Kingdom), with the provision of salient points about their care and management. Working with deer is a fascinating journey of constant learning and discovery. These remarkable animals will never fail to delight and amaze those who are lucky enough to observe and interact with them.


The table below is one possible taxonomy of the Family Cervidae. It is not intended to be the definitive case. For more exhaustive accounts of deer taxonomy, the reader is redirected to Groves and Grubb (2011), Heckeberg (2020), Mackiewicz et al. (2023) and A Guide to the Deer of the World by Charles Smith-Jones (see Table 1.1).


Table 1.1 A taxonomy of deer species.



















































































































































































































































































































































































Family Sub family Tribe Genus Species Common name Notes
Cervidae Cervinae (Old world deer) Cervini Axis annamiticus Indochina hog-deer



axis Axis (chital) Found in some UK park collections



calamianensis Calamian



kuhlii Bawean deer



porcinus Hog deer Found in few UK park collections



Cervus (elaphus group) elaphus Red deer Common in farmed, wild and park collections



pannoniensis Eastern European red deer



maral Persian red deer



corsicanus Barbary



yarkandensis Yarkand



bactrianus Bactrian



hanglu Kashmir



wallichii Tibetan shou



macneilli Sichuan shou



alashanicus Alashan



xanthopygus Manchurian wapiti



canadensis Wapiti (also termed elk) Only found in certain farms. Usually not purebred; hybridised with red deer



albirostris White-lipped (Thorold’s) deer
Cervidae Cervinae (Old world deer) Cervini Cervus (nippon group) nippon Sika Multiple subspecies. Red deer hybridisation observed



aplodontus Hokkaido sika



pulchellus Tsushima sika



hortulorum Northeastern sika



sichuanicus Sichuan sika



taiouanus Taiwanese sika



pseudaxis Vietnamese sika



Elaphurus davidianus Pére David Only found in park and zoo collections



Panolia eldii Eld’s deer



thamin Thamin



siamensis Eastern Eld’s deer



Rucervus duvaucelii Barasingha Only found in park and zoo collections. Rare red deer hybridisation



ranjitsinhi Eastern swamp deer



branderi Hard-ground barasingha



Rusa alfredi Visayan spotted deer



barandanus Mindoro deer



nigellus Mindanao mountain deer



marianna Philippine spotted deer Found in some zoo collections



timorensis Rusa deer



unicolor Sambar Found in few UK park collections
Cervidae Cervinae (Old world deer) Cervini Dama dama Fallow deer Common in park collections and in the wild



mesopotamica Persian fallow deer Some genetics in park collections


Muntiaci Muntiacus (red or Indian) muntjak Indian muntjac



vaginalis Northern red muntjac



malabaricus Southern red muntjac



aureus Himalayan muntjac



nigripes Hainan muntjac



Muntiacus (crinifrons) gongshanensis Gongshan muntjac



feae Fea’s muntjac



Muntiacus (new) vuquangensis Giant muntjac



rooseveltorum Roosevelt’s muntjac



putaoensis Leaf muntjac



truongsonensis Truong Son muntjac



puhoatensis Pu Hoat muntjac



Muntiacus (reevesi group) reevesi Reeves’ muntjac Wild and park collections, not exhibition species



atherodes Bornean yellow muntjac



montanus Sumatran muntjac



Elaphodus cephalophus Tufted deer

Capreolini (New world deer) Alceini Alces alces Moose Moose in North America; in Eurasia termed elk (not to be confused with the wapiti)


Capreolini Capreolus capreolus Roe deer (western) Wild only, found in parks and not curated



pygargus Roe deer (Siberian)
Cervidae Capreolini (New world deer) Odocoileini Hydropotes Inermis (Chinese) Water deer Wild and park collections, not an exhibition species



Rangifer tarandus Reindeer (caribou) Domestic collections only



Odocoileus hemionus Mule deer



virginianus White-tailed deer



pandora Yucatan brown brocket



Blastocerus dichotomus Marsh deer



Ozotoceros bezoarticus Pampas deer



Hippocamelus antisensis Taruca



bisulcus South Andean deer (Huemul)



Mazama americana American red brocket



whitelyi Southern red brocket



gouazoubira Grey brocket



nemorivaga Amazonian brown brocket



temama Central American red brocket



bororo Small red brocket



chunyi Dwarf brocket



nana Pygmy brocket



bricenii Merida brocket



rufina Little red brocket



Pudu mephistophiles Northern pudu



puda Southern pudu

In the United Kingdom – Domestic only; Farmed only; Park only and Wild and mixed.


Axis







































Scientific name Axis axis
Male terminology Stag
Female terminology Hind
Offspring terminology Calf
Average height at shoulder 86–92 cm (M), 70 cm (F)
Average weight 80–110 kg (M), 25–45 kg (F)
Average lifespan 5–10 years
Rut All year round with a peak in April and May
Gestation length 225–230 days
Birthing (calving) All year round with a peak in winter
Usual number of offspring 1 or 2
Social structure Herding species

The Axis or ‘Chital’ deer is native to India, and it is a common ‘exotic’ deer in park collections in the United Kingdom (Figures 1.1 and 1.2). Axis are preferentially grazing animals but will browse and prefer open forest landscapes.

Group of axis and fallow deer in a grassy field.

Figure 1.1 Axis deer in the foreground with four white fallow deer in the background (Peter Green).

Three axis deer on a grassy field with a forested background.

Figure 1.2 Axis deer (Jim Walsh).


They have a golden brown coat liberally smattered with white spots, with an obvious white gorget (throat) and black nose. The antlers are uncomplicated, can be impressive and usually consist of a brow tine, a long sweeping beam and a single rear top tine.


Axis can be prone to stress when handling is attempted. They are prone to panic and flee at the slightest provocation. This is further complicated by the unpredictable antler and calving cycles, meaning that no particular time of year can guarantee that there are no stags in hard antler and no young calves at foot. If handling a herd of Axis is to be attempted, the stags should be removed beforehand, either through darting or culling. Great care should be taken when handling the remaining hinds and young.


Hand-rearing of Axis deer has been accomplished successfully, although the calves tend to remain more aloof and wary of their human surrogate than other deer species. Careful planning should be taken when re-releasing these young deer back into the park herd because they are prone to extreme panic when spooked or in unfamiliar territory. Stimulations as innocuous as the unexpected alarm call of a bird can cause Axis to charge off, seemingly blindly, sometimes running into barriers such as fences or mesh gates with fatal consequences. A staged reintroduction from a smaller release pen surrounded by an opaque barrier is recommended.


Barasingha







































Scientific name Rucervus duvaucelii
Male terminology Stag
Female terminology Hind
Offspring terminology Calf
Average height at shoulder 120–124 cm
Average weight 130–150 kg
Average lifespan 15–20 years
Rut August–October
Gestation length 240–250 days
Birthing (calving) Late April–June
Usual number of offspring 1
Social structure Herding species

Although a swampland species in their native India, Barasingha are most commonly encountered in parkland or zoological collections. The name is a combination of the Hindi words for ‘twelve’ and ‘horns’, referring to the most common antler formation grown by Barasingha stags, with six points on each antler, including a distinctive split rear tine. Despite the name, it is not unusual for Barasingha stags to have more or less than the traditional dozen points. The velvet usually has a reddish colour.


An attractive deer, Barasingha can sometimes be mistaken for red deer due to their size and similar colouration (Figures 1.3 and 1.4). They are more of a golden brown than red deer in summer and the winter coat is a dull yellowy brown. The tail is short with a white underside and the ears are larger and rounder-edged than the ears of the red deer.

Two Barasingha deer walking across a grassy field with trees in the background.

Figure 1.3 Barasingha deer (Jim Walsh).

Two barasingha deer are standing opposite on a grassy field with trees in the background.

Figure 1.4 Barasingha deer (Jim Walsh).


Barasingha can be difficult to sedate and can be prone to stress during and following capture operations. They are, therefore, challenging to handle and can pose a significant risk of injury to those attempting to handle them.


Fallow Deer







































Scientific name Dama dama
Male terminology Buck
Female terminology Doe
Offspring terminology Fawn
Average height at shoulder 85–95 cm (M); 75–85 cm (F)
Average weight 60–100 kg (M); 30–50 kg (F)
Average lifespan 12 years
Rut September–November
Gestation length 225–230 days
Birthing (fawning) June–July
Usual number of offspring 1
Social structure Herding species

The current northern European population of fallow deer is thought to be descended mostly from animals introduced by the Romans and then later by the Normans for hunting and sport. They are a popular choice of species in deer parks and are established in the wild in many countries where they can form large roaming herds (Figures 1.5 and 1.6a), sometimes numbering in the hundreds.

Leucistic fallow buck with dark antlers grazing in a grassy field.

Figure 1.5 A leucistic fallow buck with a recent injury associated with rutting with penetration of the left eye (Peter Green).

Herd of fallow does and fawns in a grassy field with a woodland background.

Figure 1.6a Fallow does and fawns (including common, melanistic and menil with a few leucistic). (Peter Green).

Herd of fallow does and fawns in a grassy field with a woodland background.

Figure 1.6b Fallow bucks – common, melanistic and menil colour forms. (Peter Green).


Fallow deer come in four colour variants.



  • Common: Common fallow deer are a dull grey-brown colour during the winter, while their summer coat is a rich brown colour covered in white spots along the back and flanks. A continuous white line runs the length of the flanks, and the underside of the deer is white. A black line runs along the spine, and the rump of a common fallow deer is white with a characteristic black upside-down horseshoe shape running across the top of the rump.
  • Menil: Menil fallow deer have similar summer coat markings to those of the common colour variant; however, they are much lighter. The base coat colour is a light fawn, and the black markings along the back and tail are a light brown colour. They also possess white spots along the back and flanks.
  • Leucistic (White or Cremello): True albinism is rare in fallow deer, while white colour variants are quite common and were probably selectively bred for within parks historically. White fallow are often desirable for display herds and can also be of use when trying to locate groups of fallow within landscapes because they are easier for the human eye to pick out than common or menil animals. Fawns start life with a ginger-coloured coat that lightens with age.
  • Melanistic (Black): Melanistic fallow have a much darker coat than the other colour variants; even so, they are not truly jet black. They have black backs and tails with slightly lighter legs and undercarriages; they still have spots (in the same manner as common and menil fallow), which are very slightly lighter than the rest of the coat, but are often indistinguishable at range. The winter coat is duller and browner and still much darker than that of the other colour variants.

Herds, both wild and emparked, may be of a single colour morph or may contain a combination of different colours. Fallow can be extremely well camouflaged within their habitat, and it is not uncommon for deer managers to have difficulty locating herds even within relatively small, enclosed areas.


A major distinguishing feature of fallow deer from most other species is the structure of their antlers (see Figure 1.7). Fallow deer have ‘palmate’ antlers, where the upper part of the antler is broad and flattened like the palm of a hand. Fallow have a prominent brow tine jutting forwards just above the coronet; they often lack a bez tine and have a small trez tine; the antler then flares out into the broad palm, which can have many smaller points or ‘spellers’ coming off the rear of the antler. The antlers of fallow deer, as with all deer, generally become larger and more complex with age until the animal is in its prime. Yearling fallow bucks or ‘prickets’ will often have simple spikes of antler with little to no branching or palmation.

Fallow deer antler diagram labeled with pedicle, coronet or burr, beam, brow tine, vestigial bay tine, tray tine, palm, and spellers.

Figure 1.7 Fallow deer antler (Peter Green).


Fallow deer form herds ranging from a dozen or so animals to roving herds hundreds strong. They will both graze and browse. Woodlands frequented by fallow can be markedly reduced in the understory and have a sharp ruler-straight browse line. Fallow deer will stand on their hind limbs to reach food, so the browse line can be up to 1.7 m. Large herds of fallow can present problems to landowners and cause substantial amounts of damage to arable crops by consuming, trampling or lying on them. Out of the breeding season, the different sexes will often form smaller herds, with does and young running together as a group while the bucks form a ‘bachelor’ herd. During the rut, fallow bucks will establish rutting stands or ‘leks’, which they will defend aggressively and from which they will display to assert dominance over other males and to attract does for mating. The belching bellow of a fallow buck is an integral part of the autumnal soundscape in many places.


Fallow deer can be wary animals that stress and panic easily, making them difficult to handle. When trying to approach groups of fallow for live capture, there will often be a dominant animal, usually a lead doe, to which the rest of the group will look and follow. Identifying this animal early on can be advantageous. Removal of this animal can sometimes help capture operations as it can reduce the chance of the herd being led away from the catch area by the doe, but equally can lead to confusion and more erratic behaviour in a herd temporarily bereft of leadership.


The approach to groups of fallow deer should take into consideration the usual environment in which the particular animals exist. In wild or semi-wild populations, successful attempts to dart an animal will require careful and skilled stalking. However, slow stalking of park fallow, which are used by members of the public walking by, will instantly arouse suspicion. In such herds, boldly striding along an established footpath may be a much more effective way of approaching a group.


Fallow can be readily habituated to food, and this is often used within parks to allow health checks and close observations to be carried out. Feeding can also be used to facilitate catching or easier darting of fallow deer. Despite their size, fallow deer are immensely powerful and athletic. The hooves of fallow deer are surprisingly sharp and when cornered and not properly handled can become dangerous weapons. Handling of fallow bucks in hard antler should be avoided. When pressurised, bucks will attack humans or other deer, potentially fatally. Prickets can be aggressive to other deer, including other deer species.


When run into an appropriately designed catching and handling system with experienced handlers, fallow does and fawns settle surprisingly well and can be handled with relative ease and minimal stress. This requires experience, expertise and preparation in order to be achieved safely and humanely. All catching activities of parks or wild deer in the United Kingdom require appropriate licensing from the relevant government body.


Orphaned fallow fawns will readily adapt to hand-rearing, although the same rules that apply to the hand-rearing of buck fawns also apply to red stag calves. It should not be undertaken because of the danger a hand-reared buck can pose to human safety after puberty.


Mule Deer

































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Mar 15, 2026 | Posted by in EQUINE MEDICINE | Comments Off on Introduction to Cervids

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Scientific name Odocoileus hemionus
Male terminology Buck
Female terminology Doe
Offspring terminology Fawn
Average height at shoulder 80–106 cm
Average weight 50–150 kg (M); 43–90 kg (F)
Average lifespan 15–20 years
Rut Early November–Late December
Gestation length 200–210 days
Birthing (calving) Late May–June